Basidia transversely septate; teleutospores present; parasites.
"The New Gresham Encyclopedia. Vol. 1 Part 3" by Various
Before the teleutospore reaches maturity the nuclei fuse, and the uninucleate condition then continues again until aecidium formation.
"Encyclopaedia Britannica, 11th Edition, Volume 11, Slice 3" by Various
The teleutospore stage of many Uredineae also occurs in deep brown spots.
"Disease in Plants" by H. Marshall Ward